Carter was part of the Jamaican quartet that won the 4x100m in Beijing in 2008.

His was one of 454 selected doping samples retested by the International Olympic Committee (IOC) last year, and has been found to contain the banned stimulant methylhexaneamine.

Bolt, 30, completed an unprecedented 'triple triple' in Rio last summer.

He won gold in the 100m, 200m and 4x100m relay to add to his successes in the same events in 2008 and 2012.

Carter, 31, was also part of the squad that won the event in London five years ago and helped Jamaica win at the World Championships in 2011, 2013 and 2015.

He ran the first leg for Jamaica's 4x100m relay team in Beijing, which also included Michael Frater, Asafa Powell and Bolt.

The team won in a then-world record of 37.10 seconds, ahead of Trinidad and Tobago and Japan, who could have their medals upgraded. Brazil would then receive bronze.

The head of the Jamaica Athletics Administrative Association, Dr Warren Blake, said he did not expect the whole team to be penalised: "I didn't rule out he'd be found guilty but my personal opinion is that I'm surprised they'd go that route."

Carter's lawyer has confirmed that the sprinter will lodge an appeal with the Court of Arbitration for Sport.